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Artematopus, Artematopus, Electribius, Bruchidae, Rhaebinae, Rhaebus, Pachymerinae, Pachymerini, Pachymerus, Butiobruchus, Caryobruchus, Caryoborus, Caryedoninbi, Mimocaryedon, Caryedon, Caryotrypes, Aforedon, Caryopemonini, Exoctenophorus, Protocaryopemon, Diedobruchus, Caryopemon, Amblycerinae, Amblycerus, Amblycerini, Spermophagini, Zabrotes, Spermophagus, Eubaptinae, Eubaptus, Kytorhininae, Kytorhinus, Bruchinae, Bruchini, Bruchus, Megacerini, Megacerus, Acanthoscelidini, Gibbobruchus, Ctenocolum, Caryedes, Meibomeus, Penthobruchus, Pygiopachymerus, Merobruchus, Acanthoscelides, Mimosestes, Stylantheus, Julodinae, Julodini, Aaata, Amblysterna, Julodella, Julodis, Neojulodis, Sternocera, Polycestinae, Acmaeoderioid, Haplostethini, Ankareus, Exaesthetus, Helferella, Mastogenius, Micrasta, Namibogenius, Neomastogenius, Siamastogenius, Trigonogya, Acmaeoderini, Odetteina, Odettea, Nothomorphina, Nothomorpha, Nothomorphoides, Paracmaeoderoides, Acmaeoderoidina, Acmaeoderoides, 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Strandietta, Strigulioides, Suarezia, Toxoscelus, Xenita, Geraliina, Geralius, Pseudokamosia, Sambomorpha, Dismorphina, Cyphothorax, Dismorpha, Eudiadora, Trypantius, Amorphosomina, Amorphosoma, Asymades, Coraebina, Cryptomorpha, Dinocoroebus, Euchroaria, Holmerika, Katonia, Metasambus, Neocoraebus, Obenbergerula, Therybuprestis, Vanroonia, Coraebina, Belgaumia, Brachycoraebus, Cisseicoraebus, Coraebosoma, Coraebus, Eupristocerus, Mundaria, Philocoroebus, Meliboeina, Bourgoinia, Kamosia, Kamosiella, Laktonia, Meliacanthus, Meliboeoides, Melibaeopsis, Meliboeus, Nalanda, Tonkinula, Xenomerius, Synechocerina, Synechocera, Clemina, Anodontodora, Clema, Lepidoclema, Niehuisia, Promeliboeus, Pseudoclema, Dinocephalia, Meliboeithon, Paracephala, Agrilini, Deyrollius, Eumerophilus, Lepismadora, Nickerleola, Parasambus, Pseudagrilodes, Pseudagrilus, Sambus, Sarawakita, Wendleria, Weyersiella, Amorphosternina, Amorphosternoides, Amorphosternus, Bergidora, Diadora, Diadorina, Helferina, Amyiina, Amyia, Euamyia, Pareumerus, Rhaeboscelidina, Paragrilus, Rhaeboscelis, Velutia, Agrilina, Agrilochyseus, Agrilodia, Agriloides, Agrilus, Curtis, Autarcontes, Bellamyus, Callipyndax, Malawiella, Maublancia, Mychommatus, Omochyseus, Parakamosia, Pilotrulleum, Sjoestedtius, Aphanisticini, Aphanisticina, Aphanisticus, Endelus, Anthaxomorphina, Anthaxomorphus, Cylindromorphina, Afrocylindromorphus, Cantonius, Cylindromorphus, Franchetia, Heromorphus, Paracylindromorphus, Zitella, Zitoides, Zita, Cylindromorphoidina, Cylindromorphoides, Eurynodes, Germaricina, Germarica, Trachini, Subtribe, Habroloma, Neotrachys, Trachys, Brachina, Brachys, Lius, Taphrocerus, Leiopleurina, Callimicra, Leiopleura, Pachyschelina, Hylaeogena, Pachyschelus, Aptinus, Brachinus, Mastax, Pheropsophina, Carabinae, Calosoma, Carabus, Cychrus, Leistus, Nebria, Notiophilus, Pelophila, Cicindelinae, Cicindela, Cylindera, Elaphrinae, Blethisa, Diacheila, Elaphrus, Harpalinae, Abax, Acupalpus, Agonum, Amara, Anchomenus, 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Lorocera, Mastax, F, Metabletus, Molops, Odontonyx, Parophonus, Penthus, Polystichus, Pseudaphaenops, Taphoxenus, Thalassophilus, Trachypachys, Trichocellus, Trichotichnus, Zuphium,’ Acapnolymma, Aedoeus, Afroartelida, Anthribola, Antigenes, Apatophysis, Apheledes, Apiocephalus, Appedesis, Ariastes, Artelida, Barossus, Boppeus, Borneophysis, Capetoxotus, Capnolymma, Catalanotoxotus, Criocerinus, Dorcasomus, Dorcianus, Dotoramades, Dysmathosoma, Eccrisis, Echarista, Enthymius, Gaurotinus, Harimius, Icariotis, Kudekanye, Lepturastra, Lingoria, Logisticus, Lycosomus, Mastododera, Megasticus, Musius, Myiodola, Otteissa, Pachysticus, Paralogisticus, Paratophysis, Paratoxotus, Peithona, Phithryonus, Planisticus, Pseudogenes, Pyllotodes, Raharizonina, Ramodatodes, Rhagiops, Sagridola, Scariates, Scopanta, Stenotsivoka, Stenoxotus, Suzelia, Tomobrachyta, Toxitiades, Trichroa, Tsivoka, Villiersicus, Xanthopiodus, Agaleptoides, Agaleptus, Amblyontium, Anexamita, Anubis, Aphrodisium, Aromia, Aromiella, Asmedia, Beaveriella, Bradycnemis, Callichroma, Callixanthospila, Cataphrodisium, Chelidonium, Chloridolum, Chromalizus, Clavomela, Cloniophorus, Closteromerus, Cnemidochroma, Compsomera, Conamblys, Cotychroma, Dictator, Diotecnon, Dubianella, Embrikstrandia, Eugoa, Eugoides, Eulitopus, Euporus, Gauresthes, Gestriana, Guitelia, Helymaeus, Hospes, Huedepohliana, Hybunca, Hylomela, Hypargyra, Hypatium, Hypocrites, Ipothalia, Jonthodes, Jonthodina, Leptosiella, Linsleychroma, Litomeces, Litopus, Mattania, Mecosaspis, Metallichroma, Mionochroma, Monnechroma, Niraeus, Oligosmerus, Osphranteria, Oxyprosopus, Pachyteria, Paraguitelia, Parandrocephalus, Philematium, Philomeces, Phrosyne, Phyllocnema, Phyllomaeus, Plinthocoelium, Polyzonus, Promeces, Psephania, Psilomastix, Quettania, Rhopalizus, Rhopalomeces, Scalenus, Schmidtiana, Schmidtianum, Schwarzerion, Schwarzerium, Synaptola, Tarsotropidus, Thompsoniana, Turkaromia, Vittatocrites, Xystochroma, Zonopterus, Afromolorchus, Anencyrus, Anomoderus, Brachypteroma, Earinis, Epania, Gastrosarus, Glaphyra, Leptepania, Mecynopus, Merionoedopsis, Molorchoepania, Molorchus, Nadezhdiana, Nathrioglaphyra, Omotes, Paranomoderus, Agallissus, Osmopleura, Zagymnus, Curiosa, Curius, Marauna., —, Nyctonympha[, ], Tmesisternus[, ], Mimasyngenes[, ], Gorybia, Haruspicina, Piezocerina[, ], Pothyne[, ], Morimonella[, ], Moechotypa, Xylariopsis, Pogonocherus, Mesosa, Hylotrupes, Apomecyna, Asaperda, Microlera, Acalolepta, Anoplophora, Astynoscelis, Monochamus, Bostrychopsebium, Bottegia, Chorotyse, Cleptopsebium, Duffyia, Frondipedia, Haplopsebium, Hovorea, Idiopsebium, Macropsebium, Nathriobrium, Nathrius, Paraleptidea, Pembius, Plectopsebium, Psebium, Pseudobottegia, Cagosima, Eumecocera, Epiglenea, Eutetrapha, Mandibularia, Menesia, Mimocagosima, Paraglenea, Paramenesia, Parastenostola, Saperda, Stenostola, Thyestilla, Conizonia, Mallosia, Mallosiola, Micromallosia, Nupserha, Oberea, Oxylia, Paramallosia, Phytoecia, 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Sphaerioeme, Sphaerion, Sphaerionillum, Steata, Stenelaphus, Stenosphenus, Stizocera, Terpnissa, Thoris, Trichophoroides, Tropimerus, Urorcites, Yorkeica, Eodorcadion, Neodorcadion, Politodorcadion, Aesiotyche, Africophanes, Alastos, Anatinomma, Anoplomerus, Aspitus, Austranoplium, Austrophanes, Blabinotus, Bouchardius, Brittonella, Brothylus, Cacophrissus, Catoptronotum, Cerasphorus, Cetimaique, Cidugala, Corupella, Crinarnoldius, Dubiefostola, Eburiomorpha, Eucrossus, Eusapia, Garissa, Garnierius, Gastrophacodes, Gnatholea, Grammicosum, Haplidus, Hesperanoplium, Hespereburia, Hesperophanes, Hesperophanoschema, Hesperophymatus, Heteraneflus, Hololeprus, Limernaea, Liosteburia, Malobidion, Meganoplium, Megosmidus, Nesophanes, Oebarina, Oemophanes, Oraphanes, Osmidus, Paracorupella, Paraliostola, Parandraceps, Perilasius, Phacodes, Phrynocris, Phymatioderus, Potiaete, Rusapeana, Stromatiodes, Stromatium, Swaziphanes, Thecladoris, Tippmannia, Tomentophanes, Trichoferus, Tropicophanes, 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Xixuthrus, Eurypoda, Encyclops, Leptalia, Pyrotrichus, Desmocerus, Caraphia, Centrodera, Formosotoxotus, Leptorhabdium, Noctileptura, Palaeoxylosteus, Pseudoxylosteus, Xylosteus, Acanthocinus, Acanthodoxus, Alcathous, Alcathousites, Alcidion, Alphinellus, Amniscites, Amniscus, Anisolophia, Anisopodesthes, Anisopodus, Antecrurisa, Antilleptestylus, Apteralcidion, Astyleiopus-Astylidius, Astylopsis, Atelographus, Atrypanius, Baecacanthus, Baryssinus, Beloesthes, Brevoxathres, Callipero, Calolamia, Canidia, Canidiopsis, Carpheolus, Carphina, Carphontes, Catharestes, Chaetacosta, Cicanyssodrys, Cleodoxus, Cobelura, Coenopoeus, Colobeutrypanus, Cometochus, Cosmotoma, Cosmotomidius, Cristurges, Dectes, Eleothinus, Eneodes, Erphaea, Eucharitolus, Eugamantus, Eutrichillus, Eutrypanus, Exalcidion, Exocentrus, Fisherostylus, Glaucotes, Granastyochus, Hamatastus, Hexacona, Hylettus, Hyperplatys, Idephrynus, Lagocheirus, Lasiolepturges, Leiopus, Leptostylopsis, Leptostylus, Leptrichillus, Lepturdrys, Lepturgantes, Lepturges, Lepturgotrichona, Liopinus, Lithargyrus, Maculurges, Mecotetartus, Nealcidion, Neoeutrypanus, Neopalame, Neseuterpia, Nyssocarinus, Nyssodectes, Nyssodrysina, Nyssodrysternum, Oectropsis, Oedopeza, Olenosus, Onalcidion, Oxathres, Ozineus, Palame, Paranisopodus, Paratenthras, Paroecus, Pattalinus, Pentheochaetes, Phrissolaus, Piezochaerus, Probatius, Proxatrypanius, Pseudastylopsis, Pseudocobelura, Pseudolepturges, Pucallpa, Sciadosurus, Sphenopsilus, Sporetus, Stenolis, Stenopsilus, Sternidius, Sternidocinus, Styloleptus, Sympagus, Tenthras, Tithonus, Toronaeus, Trichalphus, Trichastylopsis, Trichocanonura, Trichotithonus, Tropidozineus, Trypanidius, Urgleptes, Urographis, Valenus, Xenostylus, Xylergates, Aeolesthes, Allodisus, Amphelictus, Atiaia, Bolbotritus, Bothrocerambyx, Butherium, Calocerambyx, Calpazia, Cerambolbus, Cerambyx, Cevaeria, Coelodon, Coelodoniella, Coleoxestia, Criodion, Cyriopalus, Derolus, Derolydnus, Dialeges, Dissopachys, Dymasius, Elydnus, Gibbocerambyx, Hamaticherus, Hirtobrasilianus, Hoplocerambyx, Imbrius, Ischionorox, Jebusaea, Juiaparus, Jupoata, Lachnopterus, Macrambyx, Margites, Masatopes, Massicus, Melathemma, Metacriodion, Micrambyx, Nadezhdiella, Neocerambyx, Neoplocaederus, Ochrodion, Opsamates, Pachydissus, Paracriodion, Parasphallenum, Peruanus, Plocaederus, Pneumida, Poeciloxestia, Potiaxixa, Prosphilus, Pseudaeolesthes, Ptycholaemus, Rhytidodera, Sebasmia, Sphallambyx, Sphallenopsis, Sphallenum, Sphallopterus, Sphallotrichus, Tapinolachnus, Taurotagus, Teraschema, Trachylophus, Trirachys, Utopia, Xenopachys, Xestiodion, Xoanodera, Xoanotrephus, Zatrephus, Zegriades, Aegoidus, Aethecerinus, Afghanicenus, Allocerus, Allopeplus, Amannus, Amarysius, Amphionthe, Ancylosternus, Andrachydes, Andraegoidus, Anoplistes, Athetesis, Axestoleus, Batus, Batyle, Bunothorax, Calchaenesthes, Callona, Ceragenia, Cervilissa, Charinotes, Chemsakia, Chevrolatella, Chlorotherion, Chydarteres, Cosmocerus, Crioprosopus, Crossidius, Cryptobias, Ctenodes, Cyphosterna, Deltaspis, Deretrachys, Desmoderus, Dicelosternus, Dicranoderes, Dorcadocerus, Drychateres, Eleanor, Elytroleptus, Entomosterna, Eremoceras, Eriocharis, Eriphus, Erythroclea, Euryclelia, Euryphagus, Falsanoplistes, Galissus, Gambria, Georgiana, Giesbertia, Gonyacantha, Gortonia, Hoegea, Ischnocnemis, Linsleyella, Lissonoschema, Lissonotypus, Lophalia, Mannophorus, Martinsellus, Megaderus, Metaleptus, Metopocoilus, Micropelta, Molitones, Monneellus, Muscidora, Neochrysoprasis, Neocrossidius, Neogalissus, Neotaphos, Neotaranomis, Noguerana, Nothoprodontia, Oxymerus, Ozodera, Palaeotrachyderes, Panchylissus, Parabatyle, Paraethecerus, Paragortonia, Parathetesis, Parevander, Paroxoplus, Parozodera, Pavieia, Perarthrus, Phaedinus, Phimosia, Phoenicus, Phoenidnus, Pleuromenus, Plionoma, Poecilopeplus, Polyschisis, Prodontia, Pseudodeltaspis, Pseudoeriphus, Pseudophimosia, Pseudostenaspis, Pteracantha, Pteroplatidius, Pujolia, Purpuricenus, Rachidion, Retrachydes, Rhodoleptus, Schizax, Scythroleus, Seabraellus, Seabraia, Seabriella, Sphaenothecus, Steinheilia, Stenaspis, Stenobatyle, Sternacanthus, Stiphilus, Streptolabis, Tamenes, Trachelissa, Trachyderes, Trachyderomorpha, Tragidion, Triacetelus, Tuberorachidion, Tylosis, Weyrauchia, Xylocaris, Zalophia, Zenochloris, Zonotylus, Afroeme, Afromethia, Amimes, Amphelissoeme, Amplilygrus, Antennoeme, Aponoeme, Argentinoeme, Atenizoides, Atenizus, Austroeme, Calybistum, Capegaster, Catoeme, Chromoeme, Cilioeme, Coleomethia, Coptoeme, Corioeme, Cyanomethia, Elegantometallyra, Enicoeme, Etiosaphanus, Eudistenia, Euryprosopus, Gabunsaphanidus, Gounelleoeme, Graecoeme, Haplidoeme, Hyphus, Kamuia, Kenyoeme, Leptoeme, Leptoxenus, Liberedaxia, Listrocerum, Lygrus, Macroeme, Malacopterus, Martinsia, Metallyra, Methia, Methicula, Methioeme, Methioides, Mimoeme, Namiboeme, Nastocerus, Necydalosaurus, Neoeme, Neolygrus, Neomarius, Nesoeme, Noserius, Nosoeme, Nungena, Ocroeme, Oeme, Oemospila, Oplatocera, Paracalybistum, Parahyphus, Paramartinsia, Paranoplium, Paratemnopis, Paratessaropa, Parauxesis, Paroeme, Paulianometallyra, Phrynoeme, Placoeme, Proeme, Psathyrioides, Pseudomethia, Saphanidus, Senorius, Sepaicutea, Sphagoeme, Sphalloeme, Stenocoptoeme, Stenoeme, Styloxus, Temnopis, Tessaropa, Tetraommatus, Tibestia, Tristachycera, Vandykea, Xanthoeme, Xystrocera, Beraba, Cupanoscelis, Dioridium, Eburella, Eburia, Eburiaca, Eburodacrys, Eburodacrystola, Eburostola, Erosida, Neoeburia, Opades, Pantomallus, Pronuba, Quiacaua, Solangella, Styliceps, Susuacanga, Tumiditarsus, Uncieburia, Volxemia, Atelopteryx, Callisphyris, Cauarana, Hephaestioides, Hephaestion, Mendesina, Necydalis, Parahephaestion, Planopus, Platynocera, Psebena, Rhathymoscelis, Ulochaetes, Anatolobrium, Anisogaster, Bolivarita, Calobrium, Capezoum, Capobrium, Cleistimum, Comusia, Conobrium, Duffyoemida, Falsobrium, Hypomares, Ibidiomimus, Ibidionidum, Idobrium, Iphra, Iranobrium, Longipalpus, Mythozoum, Obrioclytus, Obrium, Oculobrium, Oemida, Ossibia, Oxilus, Pseudiphra, Pseudobrium, Spiniphra, Stenhomalus, Transvaalobrium, Uenobrium, Yemenobrium, Zimbaobrium, Acutelinopteridius, Ambagous, Araeotis, Aruama, Axinopalpis, Caribbomerus, Gracilia, Hybometopia, Hypexilis, Lianema, Linopodius, Linopteridius, Linopteropsis, Longelinopodius, Lucasianus, Parommidion, Penichroa, Perigracilia, Tricholinopteridius, Vesperella, Acanthonessa, Bomarion, Ectenessa, Ectenesseca, Ectenessidia, Eurymerus, Lissoeme, Meryeurus, Niophis, Tricheurymerus, Aethiora, Emenica, Neouracanthus, Rhinophthalmus, Scolecobrotus, Uracanthus, Acanthocinus Aegomorphus Anoplophora Astylopsis Ataxia Callimetopus Coenopaeus Cyrtinus Dectes Dorcaschema Ecyrus Eupogonius Goes Hemierana Hippopsis Hyperplatys Leptostylopsis Leptostylus Lepturges Liopinus Mecas Microgoes Moneilema Monochamus Neoptychodes Nyssodrysina Oberea Oncideres Oplosia Palausybra Plectrodera Plectrura Pogonocherus Psenocerus Pseudonemophas Saperda Spalacopsis Sybra Synaphaeta Tetraopes Urgleptes Urographis, Agada, Callidiellum, Callidium, Calydon, Dolomius, Elatotrypes, Gerdberndia, Hylotrupes, Isotrium, Leioderes, Lioderina, Meriellum, Oupyrrhidium, Palaeocallidium, Phymatodes, Physocnemum, Pnigomenus, Poecilium, Prionopsis, Pronocera, Pyrrhidium, Ropalopus, Semanotus, Teorotrium, Thrichocalydon, Turanium, Xylocrius, Abacoclytus, Acrocyrta, Amamiclytus, Anthoboscus, Borneoclytus, Brachyclytus, Calanthemis, Calloides, Carinoclytus, Chlorophorus, Clytocera, Clytoleptus, Clytopsis, Clytosaurus, Clytus, Cyrtoclytus, Demonax, Denticerus, Dexithea, Epiclytus, Euryscelis, Glycobius, Hesperoclytus, Ischnodora, Isotomus, Kazuoclytus, Laodemonax, Mecometopus, Megacheuma, Megacyllene, Mygalobas, Neoclytus, Neoplagionotus, Ochraethes, Paraplagionotus, Perissus, Placoclytus, Placosternus, Plagionotulus, Plagionotus, Plagithmysus, Plesioclytus, Pseudosphegesthes, Psilomerus, Rhabdoclytus, Rhaphuma, Rusticoclytus, Sarosesthes, Sclethrus, Sphegoclytus, Tanyochraethes, Teratoclytus, Trichoxys, Triodoclytus, Turanoclytus, Tylcus, Xylotrechus, Acyrusa, Adrium, Anthribatus, Araespor, Arrhytmus, Bethelium, Bouakea, Caccothorax, Callidiopis, Ceresium, Conoxillus, Conoxilus, Coscinedes, Curtomerus, Dictamnia, Didymocantha, Ectinope, Elegantozoum, Examnes, Falsoibidion, Gelonaetha, Herozoum, Laniferus, Lasiophanes, Linyra, Metacopa, Minutius, Neobethelium, Nesanoplium, Notoceresium, Oemona, Oxymagis, Paphora, Paralocus, Parasalpinia, Porithea, Psylacrida, Salpinia, Sassandria, Semiope, Sidellus, Sisyrium, Stenobrium, Stenodryas, Stenygrinum, Sternangustum, Teocchius, Tethionea, Thephantes, Trinophylum, Wahn, Zarina, Tragosoma, Anaglyptus, Aphysotes, Clytoderus, Cyrtophorus, Diphyrama, Hirticlytus, Microclytus, Oligoenoplus, Paraclytus, Pempteurys, Tilloclytus, Yoshiakioclytus, Ambeodontus, Ancylodonta, Astetholea, Bardistus, Diotimana, Imerinus, Marileus, Ommidion, Parasemnus, Phlyctaenodes, Tricheops, Acmaeops, Akimerus, Brachyta, Cortodera, Dinoptera, Encyclops, Enoploderes, Evodinellus, Evodinus, Fallacia, Gaurotes, Gaurotina, Gnathacmaeops, Grammoptera, Lemula, Macropidonia, Oxymirus, Pachyta, Pachytella, Pidonia, Pseudogaurotina, Pseudosieversia, Rhagium, _, Rhamnusium, Sivana, Stenocorus, Toxotinus, Xenophyrama, Achenoderus, Acrocyrtidus, Aglaoschema, Allopeba, Callichromopsis, Caperonotus, Chaetosopus, Chariergus, Chenoderus, Chlorethe, Compsoceridius, Compsocerus, Cosmisomopsis, Dilocerus, Ethemon, Eurybatus, Evgenius, Hylorus, Maripanus, Mimochariergus, Orthostoma, Paromoeocerus, Parunxia, Rierguscha, Rosalia, Unxia, Upindauara, Alosterna, Anastrangalia, Anoplodera, Belamira, Caraphia, Cornumutila, Corymbia, Cyphonotida, Dokhtouroffia, Etorofus, Euryptera, Eustrangalis, Judolia, Judolidia, Kanekoa, Kirgizobia, Konoa, Leptura, Lepturalia, Lepturobosca, Lycochoriolaus, Megachoriolaus, Nivellia, Nustera, Oedecnema, Pachytodes, Pedostrangalia, Pseudalosterna, Pseudovadonia, Rapuzziana, Rutpela, Stenurella, Stictoleptura, Strangalia, Strangalomorpha, Vadonia, Xestoleptura, Calliprason, Demomisis, Drototelus, Simocrysa, Stenoderus, Syllitosimilis, Syllitus, Callipogon, Diploschema, Dragomiris, Dragoneutes, Gigantotrichoderes, Gnathopraxithea, Macellidiopygus, Praxithea, Psygmatocerus, Spathopygus, Thaumasus, Torneucerus, Torneutes, Torneutopsis, Xenambyx, Callia, Oxylopsebus, Eranina, Piratininga, Cacostola, Glypthaga, Hesychotypa, Hypsioma, Trachysomus, Moneilema, Glenea, Hirtaeschopalaea, Tetrops, Mesechthistatus, Palimna, Olenecamptus, Hesperandra, Parandra, Egesina, Pterolophia, Apriona, Batocera, Bocainella, Parmena, Parmenopsis, Plectrura, Aegomorphus, Anoreina, Oplosia, Pyrianoreina, Acalodegma, Acanthinodera, Ancistrotus, Chalcoprionus, Macrodontia, Agapanthia, Agapanthiola, Calamobius, Pseudocalamobius, Theophilea, Agapanthia, Agapanthiola, Calamobius, Pseudocalamobius, Theophilea, Cyrtonops, Dandamis, Megopis, Dorysthenes, Lobarthron, Mesoprionus, Microarthron, Miniprionus, Pogonarthron, Polylobarthron, Prionus, Pseudoprionus, Psilotarsus, Titanus, Casiphia, Closterus, Erythraenus, Prionoplus, Psephactus, Rhipidocerus, Sarmydus, Tsujius, Acanthomigdolus, Anoploderma, Sachalinobia, Xenoleptura, Necydalis, Ulochaetes, Brachytria, Certallum, Hadimus, Obrida, Omophaena, Pempsamacra, Pytheus, Titurius, Aegosomatini, Anacolini, Callipogonini, Calocomini, Ergatini, Eurypodini, Macrodontiini, Macrotomini, Mallaspini, Megopidini, Meroscelisini, Prionini, Solenopterini, Anisarthron, Arhopalus, Asemum, Cephalallus, Megasemum, Nothorhina, Tetropium, Acanthomigdolus, Anoploderma, Cherrocrius, Hypocephalus, Migdolus, Paramigdolus, Sypilus, Hypocephalini, Mysteriini, Mysteria, Pathocerus, Pseudopathocerus, Cyrtonops, Dandamis, Disteniini, Capnethinius, Distenia, Disteniazteca, Melegena, Micronoemia, Nericonia, Nethinius, Noemia, Novantinoe, Nupseranodes, Saphanodes, Tengius, Typodryas, Villiersicometes, Dynamostini, Aiurasyma, Dynamostes, Heteropalpini, Heteropalpus, Pseudocometes, Ciidae, Sphindociinae, Sphindocis, Ciinae, Ciini Acanthocis, Anoplocis, Apterocis, Atlantocis, Ceracis, Cis, Cisdygma, Eridaulus, Macrocis, Xestocis, Dimerocis, Cisarthron, Dichodontocis, Dimerapterocis, Diphyllocis, Dolichocis, Ennearthron, Euxestocis, Falsocis, Hadraule, Maphoca, Mapheae, Pityocis, Ennearthon, Knablia, Lipopterocis, Malacocis, Brachycis, Neoennearthron, Nipponapterocis, Nipponocis, Odontocis, Orthocis, Cis, Paraxestocis, Plesiocis, Polynesicis, Porculus, Strigocis, Sulcacis, Entypus, Entypocis, Syncosmetus, Omogocis, Trichapus, Tropicis, Wagaicis, Orophiini, Hyalocis, Octotemnus, Orophius, Orophyus, Orophinus, Paratrichapus, Rhopalodontus, Xylographus, Xylographellini, Scolytocis, Xylographella, Coccidulinae, Lithophilini, Lithophilus, Monocorynini, Mimolithophilus, Monocoryna, Coccidulini, Adoxellus, Auladoria, Botynella, Bucolinus, Bucolus, Bura, Coccidula, Cranoryssus, Empia, Epipleuria, Eupalea, Erithionyx, Eupaleoides, Geodimmockius, Hazisia, Hypoceras, Microrhizobius, Mimoscymnus, Nothocolus, Nothorhyzobius, Orbipressus, Orynipus, Paracranoryssus, Planorbata, Psorolyma, Rhyzobius, Rodatus, Stenadalia, Stenococcus, Syntona, Noviini, Anovia, Eurodolia, Novius, Rodolia, Vedalia, Poriini, Poria, Exoplectrini, Ambrocharis, Anisorhizobius, Aulis, Chapinella, Chnoodes, Coeliaria, Cyrtaulis, Dioria, Discoceras, Exoplectra, Neorhizobius, Neoryssomus, Hovaulis, Oridia, Peralda, Rhizoryssomus, Siola, Sumnius, Azyini, Azya, Pseudoazya, Cranophorini, Cassiculus, Cranophorus, Oryssomini, Oryssomus, Pseudoryssomus, Singhikalini, Coccinellinae, Singhikalia, Coccinellini, Aages, Adalia, Adalia, Adaliomorpha, Ailocaria, Alloneda, Anatis, Anisolemnia, Anisosticta, Antineda, Anegleis, Anegleis, Pseudanegleis, Aphidecta, Archegleis, Australoneda, Autotela, Bothrocalvia, Callicaria, Calvia, Anisocalvia, Calvia, Cheilomenes, Chloroneda, Cirocolla, Cleobora, Clynis, Coccinella, Acoccinella, Chelonitis, Coccinella, Neococcinella, Coccinula, Coelophora, Coleomegilla, Cycloneda, Declivitata, Docimocaria, Dysis, Egleis, Eoadalia, Eoanemia, Eriopis, Erythroneda, Eumegilla, Harmonia, Heterocaria, Heteroneda, Hippodamia, Adaliopsis, Adonia, Asemiadalia, Ceratomegilla, Hemispaerica, Hippodamia, Neoippodamia, Semiadalia, Hysia, Illeis, Anchilleis, Bielawskia, Illeis, Leptothea, Lemnia, Lemnia, Phrynocaria, Synia, Lioadalia, Macronaemia, Allonaemia, Macronaemia, Megalocaria, Megillina, Micraspis, Microcaria, Microneda, Mononeda, Mononeda, Paramicraspis, Mulsantina, Myrrha, Metamyrrha, Myrrha, Myzia, Naemia, Neda, Nedina, Neocalvia, Neoharmonia, Nesis, Oenopia, Oiocaria, Olla, Oxytella, Palaeoneda, Paranaemia, Phrynolemnia, Procula, Propylea, Pseudadonia, Pseudoenopia, Sospita, Sphaeroneda, Spiloneda, Synona, Synonycha, Xanthadalia, Halyziini, Eothea, Halyzia, Macroilleis, Metamyrrha, Microneda, Neohalyzia, Oxytella, Protothea, Psyllobora, Vibidia, Tytthaspidini, Bulaea, Isora, Tytthaspis, Barowskia, Tytthaspis, Discotomini, Discotoma, Euseladia, Pristonema, Seladia, Vodella, Scymninae, Aspidimerini, Acarinus, Aspidimerus, Cryptogonus, Pseudaspidimerus, Stethorini, Stethorus, Allostethorus, Parastethorus, Stethorus, Scymnini, Acoccidula, Aponephus, Apseudoscymnus, Axinoscymnus, Clitostethus, Cryptolaemus, Cycloscymnus, Cyrema, Didion, Horniolus, Keiscymnus, Leptoscymnus, Midus, Nephaspis, Nephus, Bipunctatus, Depressoscymnus, Geminosipho, Nephus, Parascymnus, Scymnobius, Sidis, Turboscymnus, Parasidis, Propiptus, Pseudoscymnus, Scymniscus, Scymnodes, Apolinus, Scymnodes, Scymnomorpha, Scymnus, Mimopullus, Neopullus, Orthoscymnus, Parapullus, Pullus, Scymnus, Veronicobius, Diomini, Decadiomus, Diomus, Heterodiomus, Magnodiomus, Erratodiomus, Scymnillini, Viridigloba, Zagloba, Zilus, Selvadiini, Selvadius, Hyperaspidini, Blaisdelliana, Corystes, Corystes, Diazonema, Helesius, Hyperaspidius, Hyperaspis, Hyperaspis, Tenuisvalvae, Thalassa, Menoscelis, Thalassa, Tiphysa, Brachiacanthadini, Brachiacantha, Cyra, Hinda, Pentiliini, Calloeneis, Curticornis, Pentilia, Cryptognathini, Cryptognatha, Ortaliinae, Ortaliini, Amida, Amidellus, Anortalia, Azoria, Cinachyra, Ortalia, Ortalistes, Paramida, Rhynchortalia, Scymnhova, Zenoria, Chilocorinae, Telsimiini, Hypocyrema, Telsimia, Platynaspidini, Crypticolus, Platynaspis, Chilocorini, Anisorcus, Arawana, Axion, Brumoides, Brumus, Chilocorus, Chilocorus, Trichorcus, Cladia, Curinus, Egius, Endochilus, Exochomus, Halmus, Harpasus, Orcus, Parapriasus, Phaenochilus, Priasus, Priscibrumus, Simmondsius, Trichorcus, Xanthocorus, Zagreus, Sticholotidinae, Sukunahikonini, Hikonasukuna, Orculus, Paraphellus, Cephaloscymnini, Aneaporia, Cephaloscymnus, Neaporia, Prodilis, Prodiloides, Microweiseini, Coccidophilus, Cryptoweisea, Dichaina, Diloponis, Gnathoweisea, Microcapillata, Microfreudea, Microweisea, Nipus, Pseudosmilia, Sarapidus, Stictospilus, Carinodulini, Carinodula, Carinodulina, Carinodulinka, Serangiini, Catana, Catanella, Delphastus, Microserangium, Serangiella, Serangium, Shirozuellini, Ghanius, Medamatento, Promecopharus, Sasajiella, Shirozuella, Plotinini, Ballida, Buprestodera, Catanaplotina, Haemoplotina, Paraplotina, Plotina, Protoplotina, Sphaeroplotina, Sticholotidini, Boschalis, Bucolellus, Chilocorellus, Coelolotis, Coelopterus, Filipinolotis, Glomerella, Habrolotis, Hemipharus, Jauravia, Lenasa, Lotis, Mimoserangium, Neaptera, Nelasa, Neojauravia, Neotina, Nesina, Nesolotis, Nexophallus, Paracoelopterus, Parajauravia, Paranelasa, Paranesolotis, Parinesa, Pharopsis, Pharoscymnus, Gymnopharus, Pharoscymnus, Phlyctenolotis, Semiviride, Sticholotis, Stictobura, Sulcolotis, Synonychimorpha, Trimallena, Xamerpillus, Xanthorcus, Xestolotis, Limnichopharini, Limnichopharus, Argentipilosini, Argentipilosa, Epilachninae, Epilachnini, Adira, Afidenta, Afidentula, Afissula, Afilachna, Chnootriba, Epilachna, Henosepilachna, Macrolasia, Subafissa, Subcoccinella, Toxotoma, Epivertini, Epiverta, Madaini, Bambusicola, Cynegetis, Damatula, Lorma, Mada, Malata, Megatela, Merma, Pseudodira, Tropha, Eremochilini, Eremochilus, Cucujus, Palaestes, Pediacus. Platisus, Glaphyridae, Amphicoma, Eulasia, Glaphyrus, Lichnanthe, Lichnia, Pygopleurus, Dasychaeta, Arctodium, Anthypna, Heterocerinae, Augilini, Augyles, Littorimus, Microaugyles, Centuriatus, Heterocerini, Culmus, Dampfius, Efflagitatus, Explorator, Lanternarius, Laternarius, Lapsus, Heterocerus, Neoheterocerus, Littorimus, Micilus, Peditatus, Tropicini, Tropicus, Hydraenidae, Prosthetopinae, Nucleotopini, Nucleotops, Protosthetopini, Protosthetops, Prosthetopini, Prosthetops, Pterosthetopini, Pterosthetops, Sicilicula, Parasthetops, Sebasthetops, Mesoceration, Orchymontia, Podaena, Homalaena, Hydraenidiini, Hydraenida, Haptaenida, Parthydraenida, Madagaster, Gondraena, Coelometopon, Adelphydraena, Hydraena, Hadrenya, Haenydra, Phothydraena, Pneuminion, Parhydraena, Parhydraena, Decarthrocerus, Protozantaena, Discozantaena, Laeliaena, Limnebius, Ochtheosini, Ochtheosus, Meropathina, Meropathus, Tympallopatrum, Tympanogaster, Neochthebius, Neochthebiina, Protochthebniina, Protochthebius, Enicocerina, Enicocerus, Ochthebiina, Hughleechia, Microgasma, Ochthebius, Ochthebius, Asiobates, Calobius, Gymnochthebius, Gymnanthelius, Aulacochthebius, Abaeletes, Abraeomorphus, Abraeus, Acritodes, Acritus, Acrolister, Adelopygus, Aeletes, Aeletodes, Aemulister, Afrosoma, Alienister, Alloiodites, Althanus, Ammostyphrus, Anaglymma, Anapleus, Anasynodites, Aneuterapus, Anophtaeletes, Antongilus, Aphanister, Aphelosternus, Apobletes, Apobletodes, Arbolister, Aristomorphus, Asiaster, Asolenus, Aspidolister, Asterix, Atholus, Athomalus, Atribalus, Atribalus, Attalister, Aulacosternus, Australomalus, Axelinus, Bacaniomorphus, Bacanius, Baconia, Bastactister, Blypotehus, Cachexia, Caenolister, Caerosternus, Campylorhabdus, Canarinus, Carcinops, Catacraerus, Ceratohister, Chaetabraeus, Chaetobacanius, Chaetobacanius, Chalcionellus, Chalcurgus, Cheilister, Chelonosternus, Chelyocephalus, Chelyoxenus, Chivaenius, Chlamydopsis, Chrysetaerius, Clientister, Coelister, Coelocraera, Colonides, Conchita, Contipus, Convivister, Coomanister, Coproxenus, Coptorophis, Coptosternus, Cossyphodister, Cryptomalus, Ctenophilothis, Cyclechinus, Cyclobacanius, Cylindrolister, Cylistosoma, Cypturus, Daitrosister, Daptesister, Dendrophilus, Desbordesia, Diabletes, Dimalus, Diplostix, Discoscelis, Dolicholister, Eblisia, Ebonius, Ecclisister, Ecitonister, Ectatommiphila, Enicosoma, Eopachylopus, Epiechinus, Epierus, Epierus, Epiglyptus, Epitoxasia, Epitoxus, Epuraeosoma, Eremosaprinus, Eretmotus, Euclasea, Eucurtia, Eucurtiopsis, Eudiplister, Eugrammicus, Eulomalus, Eurysister, Euspilotus, Eutriptus, Euxenister, Exaesiopus, Exorhabdus, Exosternus, Exotoxus, Fistulaster, Gallaster, Geocolus, Geomysaprinus, Glymma, Glyptosister, Gnathoncus, Gomyopsis, Grammopeplus, Halacritus, Hemicolonides, Hesperodromus, Hetaerimorphus, Hetaeriobius, Hetaeriodes, Hetaerius, Heudister, Hindophelister, Hippeutister, Hister, Histeridae, Hololepta, Homalopygus, Hubenthalia, Hypobletus, Hypocacculus, Hypocaccus, Iberacritus, Idister, Idolia, Iliotona, Inquilinister, Iugulister, Kanaarister, Kaszabister, Kissister, Kleptisister, Lactholister, Latinolister, Latronister, Leptosister, Lewisister, Liopygus, Lissosternus, Macroabraeus, Macrosternus, Margarinotus, Margarinotus, Mascarenium, Mecistostethus, Megalocraerus, Merohister, Mesostrix, Mesynodites, Metasynodites, Microlister, Monachister, Monoplius, Monotonodites, Morphetaerius, Murexus, Myrmetes, Nagelius, Neohister, Neopachylopus, Neosantalus, Nevermannister, Nikotickis, Niponius, Niposoma, Niposoma, Nomadister, Notocoelis, Notodoma, Notolister, Notosaprinus, Nunbergia, Nymphister, Oaristes, Omalodes, Omotropis, Onthophilus, Opadosister, Operclipygus, Orectoscelis, Orphistes, Oudaimosister, Oxysternus, Pachycraerus, Pachylister, Pachylomalus, Pachylopus, Pactolinus, Panoplitellus, Parabraeus, Parahypocaccus, Paramyrmetes, Paraphilothis, Parasynodites, Paratropinus, Paratropus, Paravolvulus, Parepierus, Parodites, Paroecister, Paromalus, Pelatetister, Pelorurus, Peploglyptus, Perfidolenus, Petalosoma, Pheidoliphila, Phelister, Philothis, Philoxenus, Pholioxenus, Phoxonotus, Pinaxister, Placodes, Plaesius, Plagiogramma, Plagioscelis, Platybletes, Platyeutidium, Platylister, Platylomalus, Platysoma, Platysomatinus, Plaumannister, Poneralister, Probolosternus, Procolonides, Procoryphaeus, Psalidister, Pselaphister, Pseudepierus, Pseudister, Psiloscelis, Pterotister, Pulvinister, Pygocoelis, Reichardtia, Reichardtiolus, Reninoides, Reninopsis, Reninus, Rhypochares, Saprinillus, Saprinodes, Saprinus, Sardulus, Sartrapister, Satrapes, Scaphidister, Scapicoelis, Scapolister, Scapomegas, Sculptura, Seitzister, Sigillum, Silinus, Sitalia, Spathochus, Spelaeabraeus, Spelaeacritus, Sphaericosoma, Sphyracus, Spilodiscus, Sterncoelopsis, Sternocoelis, Sternoglyphus, Stictostix, Styphrus, Symphilister, Synetister, Synoditinus, Synoditulus, Tarsilister, Teinotarsus, Terametopon, Terapus, Teratolister, Teratosoma, Termitolister, Termitoxenus, Thaumataerius, Therondus, Theropatina, Thoraxister, Tomogenius, Tribalasia, Tribalister, Triballodes, Tribalus, Trichoreninus, Troglobacanius, Troglosternus, Trypanaeus, Trypeticus, Trypobius, Tubulister, Tylois, Ulkeopsis, Ulkeus, Voratister, Vuattuoxinus, Wasmannister, Xenister, Xenonychus, Xenophilothis, Xenosternus, Xerosaprinus, Xestipyge, Xylonaeus, Xylostega, Yarmister, Yelamos, C, Zabromorphus, Zorius, Aesalus, Agnus, Aegus Aegus, Aegus Aegotypus, Aegognathus, Sclerostomus Altitatiayus, Allotopus, Aegus Alcimus, Amneidus, Aphanognathus, Apterocyclus, Apterodorcus, Serrognathus Aulacostethus, Auxicerus, Sphaenognathus Australognathus, Bartolozziolucanus, Beneshius, Bomansius, Chiasognathus Bomansodus, Brasilucanus, Serrognathus Brontodorcus, Cacostomus, Calcodes, Pycnosiphorus Calvertius, Cantharolethrus, Capreolucanus, Cardanus, Casignetus, Ceratognathus, Ceruchites, Ceruchus, Charagmophorus, Ganelius, Geodorcus, Aegus Gnaphaegus, Gnaphaloryx Gnaphaloryx, Gonometopus, Hemisodorcus Hemisodorcus, Heterochthes, Prosopocoilus Homoderinus, Hexarthrius, Holloceratognathus, Homoderus, Homolamprima, Prosopocoilus Hoplitocranum, Hoplogonus, Lamprima, Serrognathus Lasiodorcus, Lissapterus, Leptinopterus, Lissotes, Lucanobium, Lucanus Lucanus, Macrocrates, Macrodorcas Macrodorcas, Prosopocoilus Macrodorcinus, Oonotus Macroonotus, Mesotopus, Metadorcinus, Metadorcus, Metallactulus, Prosopocoilus Metopodontus, Aegus Micraegus, Microlucanus, Mitophyllus, Macrodorcas Miwanus, Ceratognathus Neoceratognathus, Neolucanus, Nicagus, Nigidionus, Nigidius, Hemisodorcus Nipponodorcus, Novonigidius, Odontolabis, Onorelucanus, Oonotus Oonotus, Gnaphaloryx Pachistaegus, Paleognathus, Hemisodorcus Paradorcus, Aegus Paraegus, Paralissotes, Macrodorcas Pelecognathus, Penichrolucanus, Phalacrognathus, Platyceroides, Platyceropsis, Platycerus, Platyfigulus, Macrodorcas Pogonodorcus, Prismognathus, Prosopocoilus Prosopocoilinus, Prosopocoilus Prosopocoilus, Protognathinus, Aegus Pseudaegus, Prosopocoilus Psalidoremus, Prosopocoilus Pseudodontolabis, Pseudodorcus, Lucanus Pseudolucanus, Pseudorhaetus, Pycnosiphorus Pycnosiphorus, Rhaetulus, Rhaetus, Rhyssonotus, Sclerostomus Sclerostominus, Sclerostomus Sclerostomulus, Sclerostomus Sclerostomus, Scortizus, Serrognathus Serrognathus, Sinodendron, Sphaenognathus Sphaenognathinus, Sclerostomus Stomosclerus, Streptocerus, Sphaenognathus Chiasognathinus, Chiasognathus Chiasognathus, Aegus Cherasphorus, Sphaenognathus Chiasornithodus, Sclerostomus Chileistomus, Prosopocoilus Cladognathinus, Prosopocoilus Cladognathus, Cladophyllus, Colophon, Cretaesalus, Cyclommatus Cyclommatellus, Cyclommatus Cyclommatinus, Cyclommatus Cyclommatus, Prosopocoilus Cyclotropus, Dendezia, Dendroblax, Macrodorcas Digonophorus, Dinonigidius, Serrognathus Ditomoderus, Dorcasoides, Dorculus, Dorcus, Hemisodorcus Durelius, Dynodorcus, Echinoaesalus, Eligmodontus, Aegus Elsion, Macrodorcas Epidorcus, Epipedesthus, Pycnosiphorus Erichius, Pycnosiphorus Epipedus, Eucarteria, Eulepidius, Serrognathus Eurytrachelus, Macrodorcas Falcicornis, Figulus, Succiniplatycerus, Sphaenognathus Sphaenognathus, Syndesus, Telodorcus, Tetrarthrius, Aegus Torynognathus, Aegus Tumidaegus, Vinsonella, Pycnosiphorus Weinreichiellus, Velutinodorcus, Weinreichius, Xiphodontus,Amecomycter, Mectemyctor, Mecomycter, Mauroniscus, Pterogeniidae, Apogenia, Histanocerus, Katagenius, Kryptogenius, Labidocera, Pterogenius, Tychogenius, Rhysodidae, Rhysodinae, Rhysoditae, Arrowina, Clinidium, Arctoclinidium, Clinidium, Mexiclinidium, Protainoa, Tainoa, Dhysores, Grouvellina, Kaveinga, Angekiva, Ingevaka, Kaveinga, Vakeinga, Kupeus, Leoglymmius, Medisores, Neodhysores, Omoglymmius, Boreoglymmius, Caeconavitia, Carinoglymmius, Hemiglymmius, Indoglymmius, Laminoglymmius, Navitia, Nitiglymmius, Omoglymmius, Orthoglymmius, Pyxiglymmius, Plesioglymmius, Ameroglymmius, Juxtaglymmius, Plesioglymmius, Rhysodes, Rhyzodiastes, Rhyzoarca, Rhyzodiastes, Rhyzostrix, Rhyzotetrops, Temoana, Shyrodes, Sloanoglymmius, Srimara, Tangarona, Xhosores, Yamatosa, Scarabaeoidea, Lamellicornia, Scarabaeidae, Aegialiinae, Aegialiini, Silluviini, Aegialia, Psammoporus, Dimalia, Caelius, Leptaegialia, Rhysothorax, Anomalaegialia, Anomalaegialia, Silluvia, Micraegialia, Saprus, Sapriniana, Eremazini, Eremazus, Millingenia, Tolisus, Aphodiinae, Aphodiini, Acanthaphodius, Anomalurobius, Aphodius, Aphodius, Cytoderhinus, Digonorhinus, Hemicyclorhinus, Aphodoharmogaster, Apsteiniella, Jacobsonia, Yacobsonia, Candezeollus, Cephalocyclus, Cnemargulus, Cnemisus, Ahermes, Cnemargus, Cnemargus, Coptochirus, Dialytellus, Dialytes, Doaphius, Euheptaulacus, Exaphodius, Harmodactylus, Heptaulacus, Heptaulaculus, Koreoxyomus, Lorditomaeus, Macroretrus, Mothon, Mozartius, Osmanius, Oxycorythus, Oxyomus, Amidorinus, Renaudius, Paracoptochirus, Psammodaphodius, Sugrames, Sybax, Hypoplatys, Turanella, Isochirus, Xenoheptaulacus, Xeropsamobeus, Didactyliini, Didactylia, Glaraphodius, Nataia, Orocanthus, Pseudomothon, Pseudoxyomus, Trigonoscelus, Proctophanini, Australaphodius, Drepanocanthus, Goiginus, Harmogaster, Odontaphodius, Proctophanes, Proctammodes, Eupariini, Afroataenius, Airapus, Anaetius, Annegialia, Aphotaenius, Arupaia, Ataenioides, Ataeniopsis, Ataenius, Auperia, Auperia, Brancoataenius, Euparia, Euparia, Hexalus, Oxyomus, Pseudoammoecius, Stenataenius, Australammoecius, Batesiana, Bobiricola, Bruchaphodius, Cartwrightia, Cnematoplatys, Euparia, Euparixia, Euparixoides, Euparotrix, Fletchmanniella, Fletchmania, Haroldiataenius, Iarupea, Iarupoides, Iguazua, Lomanoxoides, Lomanoxoides, Martineziella, Myrmecaphodius, Napoa, Nettelislasia, Nettelia, Notocaulus, Caulonotus, Fahraea, Paraplesiataenius, Parataenius, Brancoataenius, Passalaphodius, Passaliolla, Phalangochaeta, Pseudataenius, Pseudosaprosites, Saprositellus, Saprosites, Cylindropterus, Saprovisca, Selviria, Setylaides, Simogonius, Liparochirus, Lomanoxiini, Lomanoxia, Psammodiini, Aphodopsammobius, Brindalus, Diastictus, Geopsammodius, Granulopsammodius, Ingogius, Leiopsammodius, Neopsammodius, Odontopsammodius, Petrovitzius, Psammodius, Psammobius, Sicardia, Bipammosus, Tesarius, Trichiopsammobius, Rhyssemini, Afrodiastictus, Bordatius, Messyrhus, Myrhessus, Mysarus, Neorhyssemus, Neotrichiorhyssemus, Pararhyssemus, Pittinius, Platytomus, Pleurophorus, Psammorpha, Rhyssemodes, Rhyssemorphus, Rhyssemus, Boucardius, Trichiorhyssemus, Phycochini, Phycochus, Thinorycterini, Thinorhycter, Rhyparini, Hadrorhyparus, Rhyparus, Antrisis, Ryparus, Ryparus, Sybacodes, Adebrattia, Australoxenella, Bruneixenus, Danielssonia, Pseudostereomera, Stereomera, Odochilini, Odochilus, Odontolochini, Gongrolophus, Loeblietus, Odontolochus, Anodontolochus, Odontoderus, Odontolytes, Termitoderini, Termitaxis, Termitoderus, Termitodiellus, Termitodius, Corythoderini, Chaetopisthes, Corythoderus, Eurycorythoderus, Stenocorythoderus, Hemicorythoderus, Paracorythoderus, Termitopisthes, Termitotroginae, Aphodiocopris, Termitotrox, Aulonocneminae, Ankaratrotrox, Aulonocnemis, Dialytoderus, Manjarivolo, Palnia, Canthonini, Agamopus, Aleiantus, Amphistomus, Merodontus, Platyphymatia, Anachalcos, Chalconotus, Anisocanthon, Anonthobium, Aphengoecus, Apotolamprus, Aptenocanthon, Arachnodes, Aulacopris, Baloghonthobium, Bohepilissus, Boletoscapter, Byrrhidium, Elassocanthon, Caeconthobium, Cambefortantus, Canthochilum, Canthon, Coprobius, Coeloscelis, Paedohyboma, Glaphyrocanthon, Peltecanthon, Goniocanthon, Geocanthon, Nesocanthon, Boreocanthon, Canthomoechus, Trichocanthon, Canthonella, Ipselissus, Canthonidia, Canthonosoma, Homodesmius, Canthotrypes, Cephalodesmius, Circellium, Coproecus, Cryptocanthon, Deltepilissus, Deltochilum, Diorygopyx, Endroedyolus, Epactoides, Epilissus, Epirinus, Endrodyantus, Eudinopus, Stenodactylus, Falsignambia, Gyronotus, Hammondantus, Hansreia, Haroldius, Ponerotrogus, Afroharoldius, Phaedotrogus, Holocanthon, Ignambia, Janssensantus, Lambroma, Lepanus, Macropanelus, Madaphacosoma, Malagoniella, Megathopa, Glauconia, Megathoposoma, Melanocanthon, Mentophilus, Aulacium, Monoplistes, Namakwanus, Nanos, Nesovinsonia, Odontoloma, Diaglyptus, Oficanthon, Onthobium, Outenikwanus, Panelus, Paracanthon, Parachorius, Paronthobium, Peckolus, Penalus, Peyrierasantus, Phacosoma, Phacosomoides, Pseudarachnodes, Pseudignambia, Pseudocanthon, Pseudonthobium, Pseudophacosoma, Pycnopanelus, Saphobiamorpha, Saphobius, Sauvagesinella, Scybalocanthon, Scybalophagus, Pseudoepirinus, Sikorantus, Sinapisoma, Sphaerocanthon, Streblopus, Sylvicanthon, Tanzanolus, Temnoplectron, Tesserodon, Tetraechma, Vulcanocanthon, Xenocanthon, Zonocopris, Eucraniini, Anomiopsoides, Anomiopsis, Ennearabdus, Eucranium, Anomiopsis, Cyclodema, Psammotrupes, Glyphoderus, Eurysternini, Eurysternus, Aeschrotes, Gymnopleurini, Allogymnopleurus, Garreta, Gymnopleurus, Paragymnopleurus, Drepanopodus, Kheper, Mnematidium, Mnematium, Neateuchus, Neomnematium, Pachylomera, Pachysoma, Scarabaeus, Actinophorus, Madateuchus, Sceliages, Sisyphini, Neosisyphus, Nesosisyphus, Sisyphus, Coprinae, Coprini, Catharsius, Chalcocopris, Copridaspidus, Copris, Coptodactyla, Boucomontia, Dichotomius, Pinotus, Heliocopris, Holocephalus, Atrichius, Isocopris, Litocopris, Macroderes, Metacatharsius, Ontherus, Pseudocopris, Pseudopedaria, Synapsis, Homalocopris, Thyregis, Arrowianella, Xinidium, Parapinotus, Ateuchini, Anomiopus, Aphengium, Ateuchus, Choeridium, Bdelyropsis, Bdelyrus, Canthidium, Neocanthidium, Coptorhina, Delopleurus, Deltorhinum, Demarziella, Notopedaria, Hypocanthidium, Onychothecus, Paraphytus, Xynophron, Maraxes, Freyus, Pedaria, Pedaridium, Pseuduroxys, Pteronyx, Sarophorus, Scatimus, Scatonomus, Trichillum, Uroxys, Onitini, Acanthonitis, Allonitis, Anonychonitis, Aptychonitis, Bubas, Cheironitis, Gilletellus, Heteronitis, Janssensellus, Kolbeellus, Lophodonitis, Megalonitis, Neonitis, Onitis, Platyonitis, Pleuronitis, Pseudochironitis, Tropidonitis, Phanaeini, Bolbites, Coprophanaeus, Megaphanaeus, Dendropaemon, Diabroctis, Taurocopris, Gromphas, Homalotarsus, Megatharsis, Oruscatus, Oxysternon, Phanaeus, Lonchophorus, Onthurgus, Sulcophanaeus, Tetramereia, Eurypodea, Boucomontius, Oniticellini, Anoplodrepanus, Cyptochirus, Drepanocerus, Drepanoplatynus, Euoniticellus, Helictopleurus, Heterosyphus, Liatongus, Australobolbus, Blackbolbus, Blackburnium, Bolbaineus, Bolbaffer, Bolbaffroides, Bolbapium, Bolbelasmus, Bolbocaffer, Bolboceras, Bolbocerus, Bolbocerastes, Bolboceratex, Bolboceroides, Bolbocerosoma, Bolboleaus, Bolborhacium, Bolbohamatum, Bolborhinum, Bolborhombus, Bolbothyreus, Bolbotrypes, Bradycinetulus, Elephastomus, Eucanthus, Halffterobolbus, Indobolbus, Meridiobolbus, Microbolbus, Mimobolbus, Namibiobolbus, Namibiotrupes, Odonteus, Parabolbapium, Prototrupes, Pseudoathyreus, Somalobolbus, Anoplotrupes, Cretogeotrupes, Enoplotrupes, Eogeotrupes, Epigeotrupes, Geohowdenius, Geotrupeslatreille, Gilletinus, Glyptogeotrupes, Halffterius, Haplogeotrupes, Aphodius, Jekilius, Lepidoderica, Lethrus, Megatrupes, Mycotrupes, Neoathyreus, Odontotrupes, Onthotrupes, Pachypus, Peltotrupes, Athyreus, Phanaeus, Phelotrupes, Sericotrupes, Silphotrupes, Sinogeotrupes, Stenaspidius, Stereopyge, Taurocerastes, Thorectes, Trypocopris, Baraudia, Typhaeus, Zuninoeus, Ceratophyus, Ceratotrupes, Cheotrupis, Chromogeotrupes, Cnemotrupes, Geotrupidae, Rutelidae, Rutelinae, Rutelini, Catoclastus, Chlorota, Chrysophora, Ectinoplectron, Eremophygus, Fruhstorferia, Heteropelidnota, Heterosternus, Homonyx, Homothermon, Hoplopelidnota, Macraspis, Mecopelidnota, Mesomerodon, Micraspis, Oogeneius, Parachrysina, Parastasia, Parhomonyx, Parhoplognathus, Pelidnota, Pelidnota, Pelidnotopsis, Peltonotus, Peperonota, Plusiotis, Pseudogeniates, Rutela, Subpeltonotus, Xenopelidnota, Anoplognathini, Aulacopalpus, Brachysternus, Hylamorpha, Phalangogonia, Platycoelia, Anomalini, Leptohoplia, Popillia, Anomala, Geniatini, Zenogeniates, Leucothyreus, Rhizogeniates, Mimogeniates, Xenogeniates, Labogeniates, Eunanus, Geniatosoma, Heterogeniates, Evanos, Trizogeniates, Geniates, Leucothyreus, Bolax, Microchilus, Hopliini, Hoplia, Hybosoridae, Anaides, Antiochrus, Apolonychus, Araeotanypus, Callosides, Celaenochrous, Chaetodus, Coilodes, Cretanaides, Cretohybosorus, Cryptogenius, Daimothoracodes, Dicraeodon, Hapalonychoides, Hybochaetodus, Hybosoroides, Hybosorus, Hypseloderus, Kuijtenous, Liparochrus, Ranidia, Liparochrus, Metachaetodus, Microphaeolodes, Mimocoelodes, Pachyplectrus, Pantolasius, Phaeochridius, Phaeochroops, Phaeochrous, Phaeocroides, Procoilodes, Ranidichrus, Ropalichrus, Seleucosorus, Tyrannasorus, Ochodaeidae, Chaetocanthus, Chaetocanthus, Codocera, Cretochodaeus, Mioochodaeus, Namibiotalpa, Neochodaeus, Ochodaeus, Parochodaeus, Pseudochodaeus, Schizopodidae, Schizopodinae, Schizopodini, Schizopus, Dystaxiini, Dystaxia, Glyptoscelimorpha, Scydmaeninae, Eutheia, Euthia, Eutheimorphus, Euthiconus, Conoderus, Euthiodes, Ascydmus, Protoeuthia, Veraphis, Cephenniini, Cephennium, Cephennodes, Cephennomicrus, Chelonoidum, Chelonoides, Coatesia, Elacatophora, Nanophthalmus, Neseuthia, Paracephennium, Paraneseuthia, Pseudocephennium, Cyrtoscydmini, Allohoraeomorphus, Alloraphes, Anhoraeomorphus, Archiconnus, Borneosabahia, Brachycepsis, Catalinus, Chathamaenus, Euconnomorphus, Euconnus, Heteroscydmus, Homoconnus, Horaeomorphus, Vinsoniana, Leptocharis, Loeblites, Lophioderus, Madagassoconnus, Maorinus, Maoria, Microraphes, Microscydmus, Mimoscydmus, Napochomorphus, Neuraphanax, Neuraphes, Neuraphomorphus, Oneila, Parapseudoconnus, Parascydmus, Parastenichnaphes, Parastenichnus, Protandroconnus, Protoconnus, Protoscydmus, Psepharobius, Pseudoraphes, Sciacharis, Scydmoraphes, Atropidus, Stenichnaphes, Stenichnoconnus, Stenichnodes, Stenichnoteras, Stenichnus, Syndicomorphus, Syndicus, Taphroscydmus, Venezolanoconnus, Plaumanniolini, Plaumanniola, Siamitini, Siamites, Chevrolatiini, Chevrolatia, Leptoscydmini, Leptoscydmus, Scydmaenini, Adrastia, Zeemicrus, Austroscydmaenus, Ceramphis, Clavigeroscydmus, Eudesis, Palaeoscydmaenus, Pseudoeudesis, Scydmaenus, Mastiginae, Clidicini, Clidicus, Erineus, Leptochromus, Papusus, Leptomastacini, Ablepton, Leptomastax, Pyladus, Pylades, Leptomastoides, Taurablepton, Mastigini, Mastigus, Australostigus, Paleostigus, Mastigus, Stenomastigus, Scirtidae, Cyphonidae, Helodidae, Elodidae, Amplectopus, Atopida, Brachycyphon, Byrrhodes, Calvarium, Cyphanodes, Cyphanus, Cyphon, Cyphotelus, Cyprobius, Elodes, Lacodes, Hemicyphon, Exagonthus, Heterocyphon, Hydrocyphon, Indiocyphon, Macrocyphon, Macrodascillus, Macrohelodes, Memorocyphon, Mescirtes, Mesocyphon, Microcara, Ora, Paneveronatus, Parelodes, Prionocyphon, Prionoscirtes, Pseudomicrocara, Pseudoscirtes, Sacodes, Flavohelodes, Sarabandus, Scirtes, Sclerocyphon, Stenactyla, Veronatus, Hysteroidea, Sphaeritidae, Sphaerites , dimidiatus, glabratus, politus, bodemeyeri, abdominale, abyssinicum, alternans, amplum, amurense, anamalaiense, andrewesi, angolense, angustatum, antennatum, anthrax, arrowi, aterrimum, atricolor, atripenne, atripenne, atripes, atrosuturale, atrum, australe, baconi, badium, basaleCas, basilewskyi, benitense, bicinctum, bicolorCas, bifasciatum, bilineatithorax, bimaculatum, binhanum, binigronotatum, binominatum, binotatum, bipunctatum, bisbimaculatum, biseriatum, biundulatum, bolivianum, borneense, brunneo, carinense, castaneicolor, castaneum, celebense, cerasinum, chapuisi, cheesmanae, chujoi, cinnamoneum, clathratum, coerulans, comes, compressum, confusum, conjunctum, consimile, consobrinum, coomani, coronatum, coronatum, cyanellum, cyanipenne, decorsei, delatouchei, deletum, discerptum, disclusum, discoidale, disconotatum, distinctum, dohertyi, donckieri, dureli, egregium, elisabethae, ellenbergeri, elongatum, emarginatum, eraserheadi, exclamans, exornatum, fainanense, fasciatomaculatum, fasciatumCas, fascipenne, feai, femorale, flavofasciatum, flavomaculatum, formosanum, fossulatum, fraternum, fryi, fukienense, gabonicum, geniculatum, gibbosum, gounellei, gracile, grande, grandidieri, grouvellei, gurung, hageni, harmandi, hexaspilotum, holzschuhi, ifanense, im, impuncticolle, incisum, incrassatum, indicum, inimpressum, innotatum, inornatum, insulare, interruptum, irregulare, jacobsoni, japonicum, japonum, javanum, klapperichi, lafertei, laosense, lateflavum, latissimum, leleupi, lesnei, lineaticolle, lineatipes, longicolle, longipenne, longipes, longithorax, lucidum, lunatum, lunulatum, luteomaculatum, luzonicum, maculaticeps, madecassum, madurense, malaccanum, mangenoti, marginale, marginatum, martapuranum, mastersi, mayneui, medionigrum, melanogaster, melli, metallescens, mexicanumCas, minutum, monteithi, montivagum, morimotoi, multinotatum, multipunctatum, nathani, negrito, nepalense, nigripes, nigrocinctulum, nigromaculatum, nigrosuturale, nigrumCas, notaticolle, notaticolle, notatum, obliteratum, ocellatum, oculare, optabile, orbiculosum, ornatum, overlaeti, palonense, pantherinum, papuanum, pardaleCas, patinoi, peninsulare, peraffine, perpulchrum, philippense, phungi, plagatum, politum, prolongatum, pulchellum, punctaticolle, punctatumCas, punctipenne, pygidiale, quadriguttatum, quadrillum, quadrimaculatum, quadriplagatum, quadriplagiatum, quadripustulatum, quadripustulatum, quadritotatumCas, quinquemaculatum, reductum, rosenbergi, rouyeri, rubicundum, rubricolle, rubritarse, ruficolor, ruficorne, rufipenne, rufipes, rufitarse, rufofemoratum, rufonotatum, rufopygum, rufum, rugatum, salvazai, sauteri, semiflavum, semilimbatum, septemmaculatum, septemnotatum, seriatum, shelfordi, shibatai, sikorai, simileCas, simplicicolle, sinense, sinuatum, sondaicum, striatipenne, striatum, suarezicum, subdepressum, subelongatum, subpunctatum, sulcaticolle, sulcatum, sulcipennes, sunannulatum, sylhetense, takahashii, takashii, takemurai, testaceum, thakali, thomasi, thoracicum, transversale, tricolor, trimaculatum, trinotatum, tsushimense, tuberculipes, undulatum, unicolor, unifasciatum, vagefasciatum, atropygum, bicoloricolle, bicoloripes, biconjuntum, bioculatum, birmanica, effigiatum, femoratum, inconjunctum, kudatense, luluanum, medionigrum, melanopus, modigliani, multimaculatum, nigripenne, nigriventre, plagatum, portevini, prolongatum, ruficolor, rufithorax, semifasciatum, sexmaculatum, unicolor, uniplagatum, var.assamense, variabile, variegatum, vernicatum, vicinum, violaceipenne, viride, vitalisi, vittipenne, waterstradti, yasumatsui, yunnanum, Hysteroidea, Syntelidae, Syntelia, histeroides, Trachypachidae, Trachypachinae, Trachypachitae, Trachypachini, Trachypachus, Systolosomatini, Systolosoma, Notioxenus, Trogidae, Omorgus, Trox, Phoberus, Glaresis, Trictenotomidae, Autocrates, Trictenotoma, childrenibirmana, childreni, doriae, thomsoni, waldenfelsi,